A "pointer" to the governing node (head) for dependency relation at the analytical level (for more info on the analytical level, see Manual for Analytic Layer Tagging of the Prague Dependency Treebank (English translation) (or the Czech original)). It "points" to the governing node with the corresponding <r> element (textual parts - #PCDATA - of the markup must match).
Pointer to 0 (zero) is allowed, and in fact required somewhere in every sentence, even tough it is never present in any r element in the sentence; it denotes the "virtual" (or technical) extra root node of the whole sentence structure. I.e. the nodes pointing to 0 are the "linguistic" roots.
For a dependency link at the tectogrammatical level, see the <TRg> element.
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